Gomphoneis herculeana

(Ehrenberg) Cleve 1894      Category: Asymmetrical biraphid
BASIONYM: Gomphonema herculeanum Ehrenberg 1845
SYNONYM(S): Gomphoneis herculeana var. robusta  (Grunow) Cleve | Gomphonema capitatum var. herculaneum sensu H.L. Smith 1877 

Gomphoneis eriense var. variabilis


Gomphoneis herculeana var. abundans

LM scalebar = 10 µm = 20 pixels.


Contributor: Pat Kociolek | Gene Stoermer - January 2011
Length Range: 50-165 µm
Width Range: 22-34 µm
Striae in 10 µm: 8-12 in the central valve, 12-14 at the headpole, 13-16 at the footpole


Valves are robust, strongly lanceolate-clavate with a broadly rounded headpole and narrow, rounded footpole. The axial area is narrow, straight, and at midvalve forms a relatively small, irregularly rounded central area that bears a single stigma. Raphe lateral, weakly undulate, with small external proximal ends. Striae are composed of double rows of areolae, arranged alternately. From the headpole to the central nodule the striae are radiate; from the central nodule to the footpole striae are radiate, become parallel, and near the footpole striae become strongly radiate. The longitudinal lines of G. herculeana at the middle portion of the valve are positioned approximately one-half the way between the axial area and margin. The apical pore fields are distinctly bi-lobed.

Initial valves are 140-165 µm long, 26-34 µm wide.

Distribution:This species, though commonly recorded across the USA, is restricted to the upper Laurentian Great Lakes and some localities in the Pacific Northwest.

Original Description

Basionym: Gomphonema herculeanum
Author: Ehrenberg 1845
Length Range: µm
Striae in 10 µm:

Original Description

Original Images

Cite This Page:
Kociolek, P., and Stoermer, G. (2011). Gomphoneis herculeana. In Diatoms of the United States. Retrieved January 18, 2017, from http://westerndiatoms.colorado.edu/taxa/species/gomphoneis_herculeana

Species: Gomphoneis herculeana

Contributor: Pat Kociolek | Gene Stoermer

Reviewer: Sam Rushforth


Cleve, P.T. (1894). Synopsis of the Naviculoid Diatoms, Part I. Kongliga Svenska-Vetenskaps Akademiens Handlingar 26(2):1-194, 5 pls.

Kociolek, J.P. and Stoermer, E.F. (1988). Taxonomy, ultrastructure and distribution of Gomphoneis herculeana, G. eriense and closely related species (Naviculales: Gomphonemataceae). Proc. Acad. Nat. Sci. Phil. 140: 24-97. http://www.jstor.org/stable/4064935

Kociolek, J.P. and Stoermer, E.F. (1991). Taxonomy and ultrastructure of some Gomphonema and Gomphoneis taxa from the Upper Laurentian Great Lakes. Canadian Journal of Botany 69: 1557-1576. 10.1139/b91-200

Kociolek, J.P. and Stoermer, E.F. (1986). Observations on North American Gomphoneis (Bacillariophyceae). II. Descriptions and ultrastructure of two new species. Transactions of the American Microscopical Society 105: 141-151 . http://www.jstor.org/stable/3226386

Links & ID's

Index Nominum Algarum (INA)

Transfer INA

California Academy of Sciences (CAS)

Gomphoneis herculeana CAS

North American Diatom Ecological Database (NADED)

NADED ID: 36003

Autecology Discussion

EMAP Assessment

The Environmental Protection Agency (EPA) western Environmental Monitoring and Assessment Program (EMAP) study was completed during the years 2000-2004 (see citations at bottom of this page). Over 1200 streams and rivers in 12 western states (Arizona, California, Colorado, Idaho, Montana, Nevada, North Dakota, Oregon, South Dakota, Utah, Washington and Wyoming) were selected for sampling based on a stratified randomized design. This type of design insures that ecological resources are sampled in proportion to their actual geographical presence. Stratified randomized design also allows for estimates of stream length with a known confidence in several “condition classes” (good or least-disturbed, intermediately-disturbed, and poor or most-disturbed) for biotic condition, chemistry and habitat.

EMAP Distribution

Gomphoneis herculeana

EMAP Response Plots

Gomphoneis herculeana

EMAP citations

Results are published in:

Johnson, T., Hermann, K., Spaulding, S., Beyea, B., Theel, C., Sada, R., Bollman, W., Bowman, J., Larsen, A., Vining, K., Ostermiller, J., Petersen, D. Hargett, E. and Zumberge, J. (2009). An ecological assessment of USEPA Region 8 streams and rivers. U.S. Environmental Protection Agency Region 8 Report, 178 p.

Stoddard, J. L., Peck, D. V., Olsen, A. R., Larsen, D. P., Van Sickle, J., Hawkins, C. P., Hughes, R. M., Whittier, T. R., Lomnicky, G. A., Herlihy, A. T., Kaufman, P. R., Peterson, S. A., Ringold, P. L., Paulsen, S. G., and Blair, R. (2005). Environmental Monitoring and Assessment Program (EMAP) western streams and rivers statistical summary. U.S. Environmental Protection Agency Report 620/R-05/006, 1,762 p.

Stoddard, J. L., Peck, D. V., Paulsen, S. G., Van Sickle, J., Hawkins, C. P., Herlihy, A. T., Hughes, R. M., Kaufman, P. R., Larsen, D. P., Lomnicky, G. A., Olsen, A. R., Peterson, S. A., Ringold, P. L., and Whittier, T. R. (2005). An ecological assessment of western streams and rivers. U.S. Environmental Protection Agency Report 620/R-05/005, 49 p.