(Ehrenberg) Cleve 1894 Category: Asymmetrical biraphid
BASIONYM: Gomphonema herculeanum Ehrenberg 1845
SYNONYM(S): Gomphoneis herculeana var. robusta (Grunow) Cleve | Gomphonema capitatum var. herculaneum sensu H.L. Smith 1877
Valves are robust, strongly lanceolate-clavate with a broadly rounded headpole and narrow, rounded footpole. The axial area is narrow, straight, and at midvalve forms a relatively small, irregularly rounded central area that bears a single stigma. Raphe lateral, weakly undulate, with small external proximal ends. Striae are composed of double rows of areolae, arranged alternately. From the headpole to the central nodule the striae are radiate; from the central nodule to the footpole striae are radiate, become parallel, and near the footpole striae become strongly radiate. The longitudinal lines of G. herculeana at the middle portion of the valve are positioned approximately one-half the way between the axial area and margin. The apical pore fields are distinctly bi-lobed.
Initial valves are 140-165 µm long, 26-34 µm wide.
Distribution:This species, though commonly recorded across the USA, is restricted to the upper Laurentian Great Lakes and some localities in the Pacific Northwest.
Basionym: Gomphonema herculeanum
Author: Ehrenberg 1845
Length Range: µm
Striae in 10 µm:
Cite This Page:
Kociolek, P., and Stoermer, G. (2011). Gomphoneis herculeana. In Diatoms of the United States. Retrieved May 19, 2013, from http://westerndiatoms.colorado.edu/taxa/species/gomphoneis_herculeana
Species: Gomphoneis herculeana
Reviewer: Sam Rushforth
Cleve, P.T. (1894). Synopsis of the Naviculoid Diatoms, Part I. Kongliga Svenska-Vetenskaps Akademiens Handlingar 26(2):1-194, 5 pls.
Kociolek, J.P. and Stoermer, E.F. (1986). Observations on North American Gomphoneis (Bacillariophyceae). II. Descriptions and ultrastructure of two new species. Transactions of the American Microscopical Society 105: 141-151 . http://www.jstor.org/stable/3226386
Kociolek, J.P. and Stoermer, E.F. (1988). Taxonomy, ultrastructure and distribution of Gomphoneis herculeana, G. eriense and closely related species (Naviculales: Gomphonemataceae). Proc. Acad. Nat. Sci. Phil. 140: 24-97. http://www.jstor.org/stable/4064935
Kociolek, J.P. and Stoermer, E.F. (1991). Taxonomy and ultrastructure of some Gomphonema and Gomphoneis taxa from the Upper Laurentian Great Lakes. Canadian Journal of Botany 69: 1557-1576. 10.1139/b91-200
Sampling for the Environmental Protection Agency (EPA) western Environmental Monitoring and Assessment Program (EMAP) study was completed during the years 2000-2004 (see citations at bottom of this page). Streams and rivers in 12 western states (Arizona, California, Colorado, Idaho, Montana, Nevada, North Dakota, Oregon, South Dakota, Utah, Washington and Wyoming). Over 1200 sites on streams and rivers in 12 western states (Arizona, California, Colorado, Idaho, Montana, Nevada, North Dakota, Oregon, South Dakota, Utah, Washington and Wyoming) were selected for sampling based on a stratified randomized design. This type of design insures that ecological resources are sampled in proportion to their actual geographical presence. Stratified randomized design also allows for estimates of stream length with a known confidence in several “condition classes” (good or least-disturbed, intermediately-disturbed, and poor or most-disturbed) for biotic condition, chemistry and habitat.
Results are published in:
Johnson, T., Hermann, K., Spaulding, S., Beyea, B., Theel, C., Sada, R., Bollman, W., Bowman, J., Larsen, A., Vining, K., Ostermiller, J., Petersen, D. Hargett, E. and Zumberge, J. (2009). An ecological assessment of USEPA Region 8 streams and rivers. U.S. Environmental Protection Agency Region 8 Report, 178 p.
Stoddard, J. L., Peck, D. V., Olsen, A. R., Larsen, D. P., Van Sickle, J., Hawkins, C. P., Hughes, R. M., Whittier, T. R., Lomnicky, G. A., Herlihy, A. T., Kaufman, P. R., Peterson, S. A., Ringold, P. L., Paulsen, S. G., and Blair, R. (2005). Environmental Monitoring and Assessment Program (EMAP) western streams and rivers statistical summary. U.S. Environmental Protection Agency Report 620/R-05/006, 1,762 p.
Stoddard, J. L., Peck, D. V., Paulsen, S. G., Van Sickle, J., Hawkins, C. P., Herlihy, A. T., Hughes, R. M., Kaufman, P. R., Larsen, D. P., Lomnicky, G. A., Olsen, A. R., Peterson, S. A., Ringold, P. L., and Whittier, T. R. (2005). An ecological assessment of western streams and rivers. U.S. Environmental Protection Agency Report 620/R-05/005, 49 p.